Goulmimichthys

Goulmimichthys Temporal range: Turonian ~93–90 Ma PreꞒ Ꞓ O S D C P T J K Pg N ↓
Goulmimichthys gasparini from La Frontera Formation, Colombia
Scientific classification
Domain:	Eukaryota
Kingdom:	Animalia
Phylum:	Chordata
Class:	Actinopterygii
Order:	†Crossognathiformes
Family:	†Pachyrhizodontidae
Genus:	†Goulmimichthys Cavin, 1995
Species
†G. arambourgi Cavin, 1995 (type) †G. gasparini Páramo, 2001 †G. roberti Blanco & Cavin, 2003

Goulmimichthys is an extinct genus of ray-finned fishes in the family Pachyrhizodontidae. The genus, first described by Cavin in 1995,^[1][2] is known from various Turonian age formations. The type species G. arambourgi from the Akrabou Formation in the El Rachidia Province of Morocco,^[3] and other fossils described are G. gasparini of the La Frontera Formation, Colombia, and G. roberti from the Agua Nueva Formation of Mexico.

Etymology

The genus name Goulmimichthys is a combination of the Greek ichthys, "fish" and Goulmima, the city in Morocco where the holotype fossils were found. The species epithet of the type species arambourgi refers to Camille Arambourg, a French paleontologist who worked in North Africa. The species epithet gasparini, given by María Páramo, honours Argentinian paleontologist Zulma Gasparini.^[4]

Description

The anterior process of the pelvic bone of Goulmimichthys has the same length as the four central abdominal vertebrae, a typical characteristic of the genus.^[5] The species G. roberti is known from disarticulated and articulated specimens.^[6]

Distribution and habitat

Paleogeography of the Turonian G. arambourgi G. gasparini G. roberti

Fossils of the type species of Goulmimichthys have been described from the Lower Turonian Akrabou Formation near Goulmima in eastern Morocco.^[7] Other species are described from the Turonian La Frontera Formation near Yaguará, Huila, in the Eastern Ranges of the Colombian Andes,^[8] and the Agua Nueva Formation of the Lagerstätte Vallecillo in northeastern Mexico.^[9]

The fossil fishes were found in associations with other fossil genera, as in Mexico Nursallia gutturosum, Rhynchodercetis regio, Tselfatia formosa, Vallecillichthys multivertebratum,^[10] in Morocco Ghrisichthys and Enchodus,^[11] and in Colombia Pachyrhizodus etayoi and Bachea huilensis.^[8] Alongside the fossil fish assemblages, the fossils of plesiosaurs as Mauriciosaurus fernandezi and Vallecillosaurus donrobertoi in Mexico, the mosasaur Yaguarasaurus columbianus in Colombia and the basal mosasaur Tethysaurus nopcsai and turtles in Morocco were recovered.

References

Bibliography

• Alvarado Ortega, Jesús; Luis M. Garibay Romero; Alberto Blanco Piñón; Gerardo González Barba; Francisco J. Vega, and Elena Centeno García. 2006. Los peces fósiles de la Formación Mexcala (Cretácico Superior) en el Estado de Guerrero, México. Revista Brasileira de Paleontologia 9. 261–272. Accessed 2017-10-17.
• Bardet, Nathalie; Alexandra Houssaye; Jean-Claude Rage, and Xabier Pereda Suberbiola. 2008. The Cenomanian-Turonian (late Cretaceous) radiation of marine squamates (Reptilia): the role of the Mediterranean Tethys. Bulletin de la Société Géologique de France 179. 605–622. Accessed 2017-10-17.
• Blanco, Alberto, and Lionel Cavin. 2003. New Teleostei from the Agua Nueva Formation (Turonian), Vallecillo (NE Mexico). Comptes Rendus Palevol 2. 299–306. Accessed 2017-10-17.
• Castelblanco Cárdenas, Lilia Milena, and Luis Eduardo Vásquez Salamanca. 2016. GeoFlash - Boletín Semanal Institucional, 1. Servicio Geológico Colombiano. Accessed 2017-10-17.
• Cavin, Lionel; Larbi Boudad; Haiyan Tong; Emilie Läng; Jérôme Tabouelle, and Romain Vullo. 2015. Taxonomic Composition and Trophic Structure of the Continental Bony Fish Assemblage from the Early Late Cretaceous of Southeastern Morocco. PLOS One 10. e0125786. Accessed 2017-10-17.
• Cavin, Lionel. 1995. Goulmimichthys arambourgi n. g., n. sp., un Pachyrhizodontidae (Actinopterygii, Teleostei) d'une nouvelle localité à nodules fossilifères du Turonien inférieur marocain. Comptes Rendus de l'Académie des Sciences de Paris 321. 1049–1054. Accessed 2017-10-17.
• González Rodríguez, Katia A.; Luis Espinosa Arrubarrena, and Gerardo González Barba. 2013. An overview of the Mexican fossil fish record, 9–34. Mesozoic Fishes 5 – Global Diversity and Evolution. Accessed 2017-10-17.
• González Rodríguez, Katia A., and Gloria Arratia. 2010. Fifth International Meeting on Mesozoic Fishes - Global Diversity and Evolution - Abstract book and fieldguides, 1–141. Universidad Autónoma del Estado de Hidalgo. Accessed 2017-10-17.
• Ifrim, Christina. 2006. The fossil Lagerstätte at Vallecillo, north-eastern Mexico: pelagic Plattenkalks related to Cenomanian-Turonian boundary anoxia (PhD thesis), 1–151. Universität Fridericiana Karlsruhe. Accessed 2017-10-17.

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Teleostei sensu de Pinna, 1996 (= Teleosteomorpha)

Kingdom: Animalia Phylum: Chordata Class: Actinopterygii Subclass: Neopterygii
Actinopterygii see Actinopterygii Neopterygii see Neopterygii Teleostei see below↓

Teleostei
Teleostei †Atacamichthys †Baleiichthys †Barschichthys †Bunoderma †Ceramurus †Dorsetichthys †Elpistoichthys †Eurycormus †Galkinia †Grimmenichthys †Hengnania †Hulettia? †Hungkiichthys †Ichthyokentema †Jialingichthys †Koonwarria †Ligulella †Luxembourgichthys †Majokia †Marcopoloichthys †Neobaleiichthys †Oligopleurus? †Pholidophorichthys †Pholidophoroides †Pholidophoropsis †Prohalecites †Ruedersdorfia †Seefeldia †Seinstedtia †Signeuxella †Pachycormidae Asthenocormus Australopachycormus Bonnerichthys Eugnathides? Euthynotus Germanostomus Haasichthys Hypsocormus Kaykay Leedsichthys Martillichthys Notodectes Ohmdenia Orthocormus Pachycormus Protosphyraena Pseudoasthenocormus Rhinconichthys Sauropsis Saurostomus Simocormus †Aspidorhynchidae Aspidorhynchus Belonostomus Jonoichthys Pseudovinctifer Richmondichthys Vinctifer †Pholidophoridae sensu stricto Annaichthys Ceneichthys Eopholidophorus Knerichthys Lombardichthys Malingichthys Parapholidophorus Pholidoctenus Pholidophoretes Pholidophorus Pholidorhynchodon Pseudopholidoctenus Zambellichthys †Archaeomaenidae Aphnelepis Archaeomaene Gurvanichthys Oreochima Wadeichthys Zaxilepis †Siyuichthyidae Bogdaichthys Dsungarichthys Manasichthys Siyuichthys Wukangia †Pleuropholidae Austropleuropholis Gondwanapleuropholis Parapleuropholis Pleuropholis Zurupleuropholis †Ankylophoridae Ankylophorus Lehmanophorus Neopholidophoropsis Pholidophoristion Siemensichthys Songaichthys Steurbautichthys †Catervariolidae Catervariolus Kisanganichthys Songanella Teleosts with cycloid scales see below↓ Otolith taxa †Archengraulis †Bicavolithus †Doggerichthys †Sphaeronchus †Xenoleptolepis

Teleosts with cycloid scales
Teleosts with cycloid scales †Antofagastaichthys †Araripichthys †Paraclupavus †Pholidolepis †Todiltia †Leptolepidae Leptolepis Longileptolepis Proleptolepis †Luisiellidae Cavenderichthys? Luisiella Waldmanichthys? †Ascalaboidae Ascalabos Ebertichthys Tharsis †Crossognathiformes Bavarichthys Chongichthys Salminops? Varasichthyidae? Bobbichthys Domeykos Luisichthys Protoclupea Varasichthys Notelopidae Bavarichthys? Michin? Notelops Nusaviichthys Crossognathidae Apsopelix Crossognathus Pachyrhizodontidae Apricenapiscis Aquilopiscis Cavinichthys Chongichthys? Elopopsis Goulmimichthys Greenwoodella Kradimus Lebrunichthys Michin? Motlayoichthys Nardopiscis Pachyrhizodus Platinx Polcynichthys Rhacolepis Stanhopeichthys Tingitanichthys †Ichthyodectiformes Africathrissops Allothrissops Altamuraichthys Antarctithrissops Ascalabothrissops? Capassoichthys Carsothrissops Dugaldia Faugichthys Furloichthys Garganoichthys Gillicus Occithrissops Ogunichthys Pachythrissops? Prymnetes Sultanuvaisia Thrissops Verraesichthys Chuhsiungichthyidae Chuhsiungichthys Jinjuichthys Mesoclupea Bardackichthyidae Amakusaichthys? Bardackichthys Heckelichthys? Cladocyclidae Aidachar Chirocentrites Chiromystus Cladocyclus Cladocynodon Eubiodectes Ichthyodectidae Cooyoo Ghrisichthys Ichthyodectes Postredectes Xiphactinus Saurodontidae Gwawinapterus Prosaurodon Saurocephalus Saurodon Unamichthys Vallecillichthys Crown-Teleostei see below↓

Crown-Teleostei
Elopomorpha see Elopomorpha †Orthogonikleithridae Cavenderichthys? Leptolepides Orthogonikleithrus Waldmanichthys? Osteoglossomorpha see Osteoglossomorpha †Tselfatiiformes Eoplethodus Protobramidae Abisaadichthys Eusebichthys Protobrama Plethodidae Bachea Bananogmius Concavotectum Dixonanogmius Enischnorhynchus Luxilites Martinichthys Moorevillia Niobrara Paranogmius Pentanogmius Plethodus Pseudonogmius Pseudothryptodus Rhamphoichthys Syntegmodus Thryptodus Tselfatia Zanclites Clupeocephala Otocephala Euteleostei